Low Metabolism & Food Retention: The koala's lethargic nature and lazy behavior can be directly credited to its low metabolic rate. They simply don't have lots of energy to expend. Metabolism can be thought of as the rate at which organisms use energy. Because their diet is very poor, the koala's primary source of energy is extremely low. By decreasing their behavioral activity and sleeping for a majority of the day, koalas are able to conserve their energy and reduce their metabolic requirements. Low metabolic rates also allow koalas to retain food within their digestive system for a long period of time, maximizing the amount of energy able to be extracted. Cork and Warner conducted interesting studies on the digestion and metabolism of Eucalyptus foliage in koalas. Using radioisotopic markers, they examined the passage of particulate and solute digesta through the alimentary tract of the koala. They found that the solute marker was retained for longer periods of time than the particulate marker. The mean retention times for the solute and particulate markers were 213 hours and 99 hours respectively. These times are longer than those reported in most other mammals (1983). The selective retention of solutes and fine particles maximizes the energy withdrawn, particularly from non cell-wall constituents. More importantly however, the relatively quick passage of larger fibrous particles, or plant cell-wall constituents, is thought to reduce the "gut-filling" effect of the foliar diet. This extends the upper limits of food intake and ultimately increases the availability of nutrients, partially compensating for the constraints of small body size (1983). Passage of the larger fibrous particles is also beneficial because other researchers found that only 25% of the the cell-wall constituents that enter the alimentary tract are able to be digested (Cork et al., 1983). Breakdown of the cell contents is most important in the digestive process.