Venom Injection Mechanism

Elapids are proteroglyphs, or ankylopropglyphs, meaning they possess fixed, hollow 
fangs at the front of their upper jaw (maxilla) that cannot be folded against the roof of 
their mouth.  The fangs lie at the front of a relatively long maxillary bone and connect 
at the base to a duct.  The duct leads to the venom gland, which is behind the eye 
(Cogger 1992; White 1981).  The maxillary bones move relatively little during biting, 
and the fixed fangs remain erect even when the snake's mouth is shut.  Venom is 
manufactured and stored in oral glands, which consist of thick connective tissue 
surrounding a lumen.  Secretory epethelium line the lumen.  Venom is secreted 
through a duct connecting to a single fang upon contraction of a separate compressor
glanulae muscle.  The compressor muscle is attached to the outer sheath of each 
venom gland.  Fangs last 6-10 weeks and replacement fangs are located behind 
functional fang.   Because of the unique physiology of the mouth of the family Elapidae, 
typically prey are siezed and held until it has been immobilized by venom.  The 
only exception to the pattern are the Death Adders, which have a more moveable 
maxilla than other elapids, though not as movable as that of the Viperidae (Greene 1997).  
An elapid skull is displayed on the left.
Image copyright Shine, 1991.

Click here to see a digram of the architecture of the Elapid mouth.


The venom gland itself consists of two sections, a posterior main gland and an anterior secretory duct.  The duct draining the main gland passes through the accessory gland and into the hollow of the fang.  The secretory epethelia that surround the main gland secrete granules that accumulate in the lumen.  The accessory gland is lined with mucus-secreting epethelium, which serve an unknown function.  Milking of Elapid snakes give low venom yields compared to those of Viperids, mainly because Elapids have a dual compartment gland instead of a wide central lumen (Minton 1971).

Home

This page is an assignment for Biology 312, Animal Physiology, and is for educational purposes only.
email me:  schutchins@davidson.edu