Social Spacing


Roaring occurs after two red howler monkey troops confront one another.
(http://web.missouri.edu/~anthmark/redhowl.htm)

Howler monkeys reside in troops on home ranges.  These home ranges can be as small as 5 ha and as large as 60 ha, but usually they are between 10 and 30 ha(Smuts et al. 1987). Home range is very variable and varies more within a species than among the 6 different species(Smuts et al. 1987).  A study of the mantled howler monkey suggests that they travel as little as possible and are energetically constrained as a result of their low energy foliage diet (Smuts et al. 1987).  Home ranges of red howlers were observed to overlap as much as 63%.  Home ranges of mantled howlers can overlap as much as 100% (Smuts et al. 1987).  Some researchers do argue that howlers exibit territoriality despite this dramatic overlap in home range because of the intergroup interactions observed in red howler monkeys (Smuts et al. 1987).  Troop interaction is usually characterized by loud roars exchanged between 2 opposing troops.  These roars are very loud and it is suggested they are the most energetically expensive thing that howlers do (Smuts et al. 1987).  Troop displacements are rare, but one troop may be forced to retreat a small distance from the altercation site (Smuts et al. 1987).  The purpose of the dawn calls that howlers make in the early morning is to to make other troops aware of the various locations of the surrounding troops.  Eventhough howlers appear to hold claim on a certain area, they are suggested to be energetically constrained from defending an exclusive territory (Smuts et al. 1987).
    One 20-year study of mantled howlers in Costa Rica showed that the average size of home range was 27.2 ha and home range correlated most closely with the number of adults in the troop (Zucker et al. 1996).  The various troops that were studied appeared to stay on the same home ranges through the 20 year study period (Zucker et al. 1996).


This page was done as part of an assignment for Animal Behavior at Davidson College.



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