Male/Male Relationships
Males have a dominance hierarchy, which is largely determined by size (which
increases with age) (Estes 1991). When two non-musth males have an aggressive
interaction over a mate, size determines the winner. When a musth male and a
non-musth male compete over a mate, the state of musth determines the winner.
When two musth males have an aggressive interaction over a mate, size determines
the winner. Aggressive interactions rarely escalate into fights (Poole 1989).
Aggressive behaviors include ear flapping (which is also use for thermoregulation),
head jerking, head tossing, and the forward-trunk-swish. An elephant indicates
submission by writhing its trunk and lowering its head while moving backwards
and sideways (Estes 1991).
Non-musth bulls rarely have aggressive encounters. Dominance is determined by
size. Bulls will allow a larger bull preferential access to a resource, such
as a watering hole (Moss 1982). When two bulls meet that are approximately the
same size, they will interlock tusks and/or trunk bases. The bull that holds
his head higher establishes dominance. Non-musth bulls sometimes live in bachelor
herds, especially young bulls. Some bulls seem to have one or more bulls that
they have a closer association with (Estes 1991).